![賀金生[北京大學城市與環境學院教授]](/img/4/197/nBnauM3X4YTOyMDO5QDN0ETN1UTM1QDN5MjM5ADMwAjMwUzL0QzL4AzLt92YucmbvRWdo5Cd0FmLxE2LvoDc0RHa.jpg "賀金生[北京大學城市與環境學院教授]")
個人簡介
學習經歷
1984.9~1988.7, 蘭州大學生物系，獲理學學士學位（植物學）
1988.9~1991.6, 中國科學院植物研究所，獲理學碩士學位（生態學）
1996.9~1998.8, 中國科學院植物研究所，獲理學博士學位（生態學）
1999.7~2002.8, 哈佛大學進化與生物學系博士後（生物學）
2007.10~2008.1, 瑞士蘇黎世大學，The Swiss National Science Foundation Research Fellow
教育工作
1991.6~1999.11 中國科學院植物研究所，實習研究員，助理研究員，副研究員
2002.8~2008.7 北京大學生態學系，副教授
2007.1~北京大學生態學系副主任
2008.8~ 北京大學生態學系，教授
2009.3~ 西北高原生物研究所百人計畫
學術任職
《植物生態學報》編委，2004~
《生物多樣性》編委，2009~
Journal of Plant Ecology (Oxford University Press，SCI), Associate editor, 2008~
Journal of Plant Research (Springer,SCI), Editorial Board Member, 2007~2014
Frontiers in Ecology and the Environment (ESA journal,SCI), Associate editor, 2007~2014
中國植物學會植物生態學專業委員會委員，2009~
中國植物學會學術委員會委員，2009~
科學研究
研究方向
1. 植物生理生態學
2. 生態系統生態學
3. 青藏高原生態學
4. 生物多樣性的生態系統功能
當前研究領域
1、生態系統生態學：主要圍繞陸地生態系統的碳循環和氮循環, 採用實驗的手段，研究全球氣候變化影響下生態系統過程的反應。近期的研究重點是：降雨格局的改變和氮沉降如何影響溫帶草地、人工林C、N循環的主要過程。
2、植物功能屬性：通過野外植物主要生態屬性的測定，結合室內植物的解剖結構的分析，在大尺度、多生態系統類型上探討植物生態屬性的變化規律和控制因子。
3、地下生態學：通過典型生態系統地下過程的調查，探討生態系統地上和地下部分的關聯，重點探討植物地上部分得到的規律性是否可以套用到地下。
4、生物多樣性的生態系統功能：結合實驗途徑和野外調查，研究生物多樣性的生態系統功能，特別是在C, N經濟（C, N economy） 中的作用。主要的實驗系統是草原生態系統。
研究項目
1、國家重點基礎研究發展規劃（973計畫）：中國陸地生態系統碳循環及其驅動機制研究（2002-2007），專題主持
2、國家自然科學基金重大研究計畫重點項目：我國溫帶草地碳庫分布、動態與氣候控制（2003-2006），專題主持
3、國家自然科學基金委 “創新群體”：環境生物地球化學研究（2004-2006）成員
4、國家自然科學基金重大研究計畫面上項目（90411004）：溫帶草地根系生態學的實驗研究：個體、種群和功能屬性的關聯（2005-2007），主持
5、國家自然科學基金委面上項目（30670322）：青藏高原極端環境條件下植物葉功能屬性特徵及環境的修飾作用（2007-2009），主持
6、中日韓A3前瞻計畫項目（Asia 3 Foresight Program）：Quantifying and predicting terrestrial carbon sinks in East Asia: toward a network of climate change research，A3 3+2（2007-2012），參加
7、國家科技支撐計畫（2007BAC06B00）：藏北高寒草甸區生態穩定性維持技術研究與示範（專題）（2007.9-2010.12），主持
8、國家自然科學基金委面上項目（30870381）：我國主要溫帶草地優勢植物葉片、根系和土壤C：N：P化學計量學的關聯（2009-2011），主持
9、歐盟項目（BACCARA）：Biodiversity And Climate Change，A Risk Analysis（2009-2011），專題3
10、科技部國際合作（Sino-EU 0916）：Risk analysis of biodiversity change：a case study in a subtropical forest，China（2009-2012），主持
11、基金委創新群體項目（31021001）：中國陸地植被的時空格局與生態功能（2011-2013），核心成員
12、"全球變化研究國家重大科學研究計畫"項目（2010CB950600）：中國陸地生態系統碳源匯特徵及其全球意義（2010-2013），研究骨幹
13、國家傑出青年科學基金（31025005）：植物功能屬性與化學計量生態學 （2011-2014），主持
14、全球變化研究國家重大科學研究計畫項目（2014CB954004）（973）“土壤系統碳動態、機制及其對全球變化的回響”（2014-2018），首席科學家
主要榮譽
1998年，中科院地奧獎學金"一等獎, 博士論文"神農架地區米心水青岡林和銳齒槲櫟林的格局和過程"
2003年，教育部自然科學一等獎：中國陸地生態系統碳循環與植被生產力的研究，第3完成人
2004年，國家自然科學二等獎：中國陸地生態系統生產力和碳循環的研究，第5完成人
2006年，北京大學“保潔”獎
2007年，Sino-Swiss Science and Technology Cooperation Research Fellowship, Swiss Federal Institute of Technology (ETH Zurich), Swiss National Science Foundation
2007年，北京大學優秀班主任三等獎
2010年，"國家傑出青年科學基金" 獲得者
2014年，北京大學“保潔”獎
主講課程
| 課程名稱 | 性質 | 學分 | 作用 | 開設時間 |
| 普通生態學（I）：個體、種群 | 本科，專業必修 | 2 | 主講 | 2005- |
| 生態學實驗技術 | 本科，專業必修 | 2 | 主講 | 2005-2007 |
| 生態系統生態學：理論與方法 | 研究生，選修 | 2 | 主講 | 2004- |
代表論著
Jing X, Sanders NJ, Shi Y, Chu HY, Classen AT, Zhao K, Chen LT, Jiang YX, He J-S* 2015.The links between ecosystem multifunctionality and above- and belowground biodiversity are mediated by climate.Nature Communications6: 8159.
Song WM, Wang H, Wang GS, Chen LT, Jin ZN, Zhuang QL, He J-S*. 2015.Methane emissions from an alpine wetland on the Tibetan Plateau: Neglected but vital contribution of non-growing season. Journal of Geophysical Research-Biogeosciences: doi:10.1002/2015JG003043.
Jin ZN, Zhuang QL, He J-S, Zhu XD, Song WM 2015.Net exchanges of methane and carbon dioxide on the Qinghai-Tibetan Plateau from 1979 to 2100. Environmental Research Letters: doi:10.1088/1748-9326/10/8/085007.
Niu KC, He J-S, Zhang ST, Lechowicz MJ 2015.Grazing increases functional richness but not functional divergence in Tibetan alpine meadow plant communities. Biodiversity and Conservation: doi: 10.1007/s10531-015-0960-2.
Chen LT, Flynn DFB, Zhang XW, Gao XL, Lin L, Luo J, Zhao CM* 2015.Divergent patterns of foliar δ13C and δ15N in Quercus aquifolioides with an altitudinal transect on the Tibetan Plateau: an integrated study based on multiple key leaf functional traits. Journal of Plant Ecology8: 303-312.
Chen LT, Flynn DFB, Jing X, Kühn P, Scholten T, He J-S 2015.A comparison of two methods for quantifying soil organic carbon of alpine grasslands on the Tibetan Plateau. PLoS ONE10: e0126372.
Mi ZR, Huang YY, Gan HJ,Zhou WJ, Flynn DFB, He J-S* 2015.Leaf P increase outpaces leaf N in an Inner Mongolia grassland over 27 years. Biology Letters 11: 20140981.
Ding S, Xu Y, Wang Y, He Y,Hou J, Chen L, He J-S 2015.Distributions of glycerol dialkyl glycerol tetraethers in surface soils ofQinghai–Tibetan Plateau: implications of GDGT-based proxies in cold and dryregions. Biogeosciences Discussion 12: 481-513.
Zhang ZJ,Yan YJ,Tian Y, LiJS, He J-S, Tang ZY* 2015.Distribution and conservation of orchid species richness in China. Biological Conservation 181: 64-72.
Eichenberg D, Trogisch S,Huang YY, He J-S, Bruelheide H 2014.Shifts in community leaf functional traits are related to litter decompositionalong a secondary forest succession series in subtropical China. Journal of Plant Ecology DOI:10.1093/jpe/rtu021: .
Wang YH, Liu HY, Chung H,Yu LF, Mi ZR, Geng Y, Jing X, Wang SP, Zeng H, Cao GM, Zhao XQ, He J-S* 2014. Non-growing-season soilrespiration is controlled by freezing and thawing processes in the summermonsoon-dominated Tibetan alpine grassland. Global Biogeochemical Cycles28: 1081-1095.
Yang XX, Yang Y, Ji CJ,Feng T, Shi Y, Lin L, Ma JJ, He J-S*2014. Large-scale patterns of stomatal traits in Tibetan and Mongoliangrassland species. Basic and Applied Ecology 15: 122-132.
Shi Y, Wang YH, Ma YL, MaWH, Liang CZ, Flynn DFB, Schmid B, Fang JY, He J-S* 2014. Field-based observations of regional-scale, temporalvariation in net primary production in Tibetan alpine grasslands. Biogeosciences11: 2003-2016.
Baumann F*, Schmidt K,Dörfer C, He J-S, Scholten T, Kühn P2014. Pedogenesis, permafrost, substrate and topography: Plot and landscapescale interrelations of weathering processes on the central-eastern TibetanPlateau. Geoderma 226-227: 300-316.
Geng Y, Wang L, Jin DM, LiuHY, He J-S* 2014. Alpine climatealters the relationships between leaf and root morphological traits but notchemical traits. Oecologia 175: 445-455.
Jing X, Wang YH, Chung H,Mi ZR, Wang SP, Zeng H, He J-S*2014. No temperature acclimation of soil extracellular enzymes to experimentalwarming in an alpine grassland ecosystem on the Tibetan Plateau. Biogeochemistry117: 39-54.
Chen LT, Niu KC, Wu Y, GengY, Mi ZR, Flynn DFB, He J-S* 2013.UV radiation is the primary factor driving the variation in leaf phenolicsacross Chinese grasslands. Ecology and Evolution 3: 4696-4710.
Jin DM, Ma JJ, Ma WH, LiangCZ, Shi Y and He J-S* 2013. Legumesin Chinese natural grasslands: Species, biomass, and distribution. Rangeland Ecology & Management 66: 648-656.
Du EZ, Zhou Z, Li P, Hu XY,Ma YC, Wang W, Zheng CY, Zhu JX, He J-S,Fang JY* 2013. NEECF: a project of nutrient enrichment experiments in China’sforests. Journal of Plant Ecology 6: 428-435.
HeJ-S*,Muraoka H, Son Y, Fang JY 2013. Carbon patterns and processes in East Asianecosystems: multi-scale approaches (Editorial). Journal of Plant Ecology6: 323-324.
Ma YL, Geng Y, Huang YY,Shi Y, Niklaus P, Schmid B, He J-S*2013. Effect of clear-cutting silviculture on soil respiration in a subtropicalforest of China. Journal of Plant Ecology 6: 335-348.
Yu LF, Wang H, Wang GS,Song WM, Huang Y, Li S-G, Liang NS, Tang YH, He J-S* 2013. A comparison of methane emission measurements usingeddy covariance and manual and automated chamber-based techniques in TibetanPlateau alpine wetland. Environmental Pollution 181: 81-90.
Liu C, Wang XP, Wu X, DaiS, He J-S, Yin WL* 2013. Relativeeffects of phylogeny, biological characters and environments on leaf traits inshrub biomes across central Inner Mongolia, China. Journal of Plant Ecology 6: 220-231.
Jin ZN*, Zhuang QL, He J-S, Luo TX, Shi Y 2013. Phenologyshift from 1989 to 2008 on the Tibetan Plateau: an analysis with aprocess-based soil physical model and remote sensing data. Climatic Change 119: 435-449.
Shi Y, Ma YL, Ma WH, LiangCZ, Zhao XQ, Fang JY, He J-S* 2013.Large scale patterns of forage yield and quality across Chinese grasslands. Chinese Science Bulletin 58: 1187-1199.
Dorfer C*,Kuhn P,Baumann F, He J-S,Scholten T 2013. Soil organiccarbon pools and stocks in permafrost-affected soils on the Tibetan Plateau. PLoS ONE 8: e57024.
Huang N*, He J-S, Niu Z 2013. Estimating thespatial pattern of soil respiration in Tibetan alpine grasslands using LandsatTM images and MODIS data. Ecological Indicators 26: 117-125.
HeJ-S 2012.Carbon cycling of Chinese forests: From carbon storage, dynamics to models. Science China Life Science 55: 188-190.
Geng Y, Wang ZH, Liang CZ,Fang JY, Frank B, Kühn P, Scholten T, HeJ-S* 2012. Effect of geographical range size on plant functional traits andthe relationships between plant, soil and climate in Chinese grasslands. Global Ecology and Biogeography 21: 416–427.
Shi Y, Baumann F, Ma YL,Song C, Kuehn P, Scholten T, He J-S*.2012. Organic and inorganic carbon in the topsoil of the Mongolian and Tibetangrasslands: pattern, control and implications. Biogeosciences 9:2287–2299.
Geng Y,Wang YH,Yang K,WangSP,Zeng H,Baumann F, Kuehn P, Scholten T, HeJ-S* 2012. Soil Respiration in Tibetan Alpine Grasslands: BelowgroundBiomass and Soil Moisture, but Not Soil Temperature, Best Explain theLarge-Scale Patterns. PLoS ONE 7: e34968.
Ma JJ, Ji CJ, Han M, ZhangTF, Yan XD, Hu D, Zeng H, He J-S*2012. Comparative analyses of leaf anatomical characteristics of dicotyledonousspecies in Tibetan and Inner Mongolian grasslands. Science China Life Sciences 42: 158-172.
Tao S*, Wang WT, Liu WX,Zuo Q, Wang XL, Wang R, Wang B, Shen GF, Yang YH, He J-S 2011. Polycyclic aromatic hydrocarbons and organochlorinepesticides in surface soils from the Qinghai-Tibetan plateau. Journal of Environmental Monitoring 13: 175–181.
Chu ZY, Lu YJ, Chang J*,Wang M, Jiang H, He J-S, Peng CH*,Ge Y 2011. Leaf respiration/photosynthesis relationship and variation: aninvestigation of 39 woody and herbaceous species in east subtropical China. Trees-Structureand Function 25: 301-310.
Ma WH, He J-S*, Yang YH, Wang XP, Liang CZ, Anwar M, Zeng H, Fang JY,Schmid B 2010. Environmental factors co-vary with plant diversity-productivityrelationships among Chinese grassland sites. Global Ecology and Biogeography 19: 233–243.
HeJ-S*, WangX, Schmid B, Flynn DFB, Li XF, Reich PB, Fang JY 2010. Taxonomic identity,phylogeny, climate and soil fertility as drivers of leaf traits across Chinesegrassland biomes. Journal of Plant Research 123: 551-561.
Ma WH, Liu ZL, Wang ZH,Wang W, Liang CZ, Tang YH, He J-S,Fang JY* 2010. Climate change alters interannual variation of grasslandaboveground productivity: evidence from a 22-year measurement series in theInner Mongolian grassland. Journal of Plant Research 123:509-517.
Baumann F, He J-S,Schmidt K, Kühn P, Scholten T. 2009. Pedogenesis, permafrost, and soil moistureas controlling factors for soil nitrogen and carbon contents across the TibetanPlateau. Global Change Biology 15: 3001–3017
Yang YH, Fang JY, Smith P,Tang YH, Chen AP, Ji CJ, Hu HF, Rao S, Tan K, He J-S. 2009. Changes intopsoil carbon stock in the Tibetan grasslands between the 1980s and 2004. Global Change Biology 15: 2723–2729
He J-S, Wang XP, Flynn DF, WangL, Schmid B, Fang JY. 2009. Taxonomic, phylogenetic and environmental tradeoffsbetween leaf productivity and persistence. Ecology 90: 2779-2791
He J-S, Wang L, Flynn DFB, WangX, Ma W, Fang J. 2008. Leaf nitrogen: phosphorus stoichiometry across Chinesegrassland biomes. Oecologia155, 301-310
He J-S, Fang JY, Wang ZH, Guo DL,Flynn DFB, Geng Z. 2006. Stoichiometry and large-scale patterns of leaf carbonand nitrogen in the grassland biomes of China. Oecologia, 149:115-122
He J-S, Wang ZH, Wang XP, SchmidB, Zuo W, Zhou M, Zheng CY, Wang MF, Fang JY, 2006. A test of the generality ofleaf trait relationships on the Tibetan Plateau. New Phytologist, 170: 835-848
Balvanera P, Pfisterer AB,Buchmann N, He J-S, Nakashizuka T, Raffaelli D, Schmid B. 2006.Quantifying the evidence for biodiversity effects on ecosystem functioning andservices. Ecology Letters, 99:1146–1156
He J-S, Flynn DFB, Wolfe-BellinK, Fang J, Bazzaz FA. 2005. CO and nitrogen, but not populationdensity, alter the size and C/N ratio of Phytolacca americana seeds. Functional Ecology, 19: 437-444
He J-S, Wolfe-Bellin KS, SchmidB, Bazzaz FA. 2005. Density may alter diversity-productivity relationships inexperimental plant communities. Basicand Applied Ecology, 6: 505-517
He J-S, Wolfe-Bellin KS, BazzazFA. 2005. Leaf-level physiology, biomass, and reproduction of Phytolaccaamericana under conditions of elevated CO and alteredtemperature regimes. International Journal of Plant Sciences, 166: 615-622
He J-S, Bazzaz FA. 2003.Density-dependent responses of reproductive allocation to elevated atmosphericCO in Phytolacca americana. New Phytologist, 157: 229-239
He J-S, Bazzaz FA, Schmid B.2002. Interactive effects of diversity, nutrients and elevated COon experimental plant communities. Oikos,97: 337-348
